Ant of the Week: Southern Fire Ant

The first thing you notice about workers of the Southern fire ant, Solenopsis xyloni, is what fierce foragers they are.

You never see just one foraging worker. Instead, there’s almost always a teeming mass.

Even when they are collecting sweets at extrafloral nectaries, Southern fire ants show up in greater numbers than most other species.

Southern fire ants are thought to be originally from throughout the southern and western United States. They have been displaced in many areas by the imported fire ant, Solenopsis invicta, but still occur widely in the dry areas of Arizona and California.

Identification:

Solenopsis ants are relatively easy to tell from other ants because of the antennae have ten segments with a two-segmented swollen area or  “club” at the end.

Solenopsis xyloni workers vary considerably in size and color, even within colonies. The larger workers tend to have lighter-colored heads and trunks than the smaller workers. In the area around Phoenix, Arizona, the Southern fire ants seem darker than those found elsewhere.

(These workers are feeding on their favorite meal of dried cat food.)

Where Solenopsis xyloni and S. invicta overlap it is difficult to distinguish the two species. Jacobson et. al. (2006) have developed a pcr technique and guidelines for identification (see references).

To make things even more confusing, it seems that Solenopsis xyloni hybridizes with Solenopsis geminata where the two overlap.  Obviously this group is “interesting” from a taxonomic standpoint.

Foraging:
The foraging workers are often seen carrying bits of hard food or arthropod parts back to the nest. They also gather some seeds.

Wet food, like this watermelon, go straight to the crop.

Often the foraging trails around their nests are underground or partially covered, so you might not notice them until you dig into the soil or pull up a weed. Then they come boiling up seemingly out of nowhere.

When foragers cross a man-made structure, such as a walking trail, sidewalk, or tile floor, they form a dense foraging trail of numerous workers traveling in both directions.

You have to admire the ability of Southern fire ants to find, process and transport food very rapidly. Plus they seem to eat just about anything they encounter. It is no wonder the colonies can grow to a relatively large size.

Do you have fire ants where you live? Have you ever watched them gather food?

References:

Solenopsis xyloni by Dale Ward

The Navajo Ant Project has a brief review of taxonomy

Ant Web shows some of the color variation within the species

Jacobson AL, Thompson DC, Murray L, Hanson SF. (2006). Establishing guidelines to improve identification of fire ants Solenopsis xyloni and Solenopsis invicta. J Econ Entomol. 99(2): 313-22.

Trager, J. C. (1991). A revision of the fire ants, Solenopsis geminata group (Hymenoptera: Formicidae, Myrmicinae).  Journal of the New York Entomological Society. 99 :141-198.

A Different Sort of “Ant”

You don’t see one of these every day.

Of course, you can tell it is a male right off*.

This one was having a bit of trouble finding its way about.

Can you spot why?

*Only male velvet ants have wings. The males do show their wasp affiliation more than the females do.

Ant of the Week: Dorymyrmex bicolor

Dorymyrmex bicolor ants are common in Arizona, especially in open areas. In a previous post, I wrote about how to identify them.

A few weeks ago I visited an agricultural research station and found almost a monoculture of Dorymyrmex.

The first thing you notice about Dorymyrmex bicolor colonies is their neat circular mounds. (Most of these were conveniently located along the dirt roads.)

Each reflect the color of the soil beneath the surface. This one wasn’t a perfect circle.

Nor this one.

Tofilski and Ratnieks (2005) studied the mound formation of two colonies of Brazilian Dorymyrmex. They found the worker ants removing excavated material from the nest deposited their loads at the crest of the mound and beyond, preventing the material from rolling back into the entrance hole. They also indicated that workers maintained the circular shape, when one side was removed, by climbing the area with the least slope to deposit their loads.

Ratnieks noted that ants of other, larger species had difficulty climbing the mounds, thus suggesting the mounds served a protective function.

As you will see in the next few photographs, the Dorymyrmex were carrying out clumps of soil and related materials the morning I visited.

Because ants at all the mounds were so busy removing material, I began to wonder about it. Were the clods of soil and pebbles rolling into the entrance hole over night? Perhaps there had been a wind storm or vehicle disturbance?

By the way, you might recognize these ants under a different name. Prior to a revision of the genera, Dorymyrmex bicolor was known as Conomyrma bicolor. Under that name, they had a brief burst of limelight when Möglich and  Alpert (1979) found they exhibited tool use by dropping small stones on their competitors, particularly Myrmecocystus workers. Finding ants that stoned their enemies caused quite a stir, as you can imagine.

Thinking about that, I also began to wonder if perhaps Dorymyrmex bicolor worker ants also drop stones on conspecifics, that is other Dorymyrmex bicolor nests. With the name change, it is somewhat hard to track the literature. Does anyone know? Anyone studying Dorymyrmex bicolor?

If not, perhaps it would be worthwhile to visit these fascinating ants some evening.

References:

Adam Tofilski and Francis L. W. Ratnieks. (2005). Sand Pile Formation in Dorymyrmex Ants. Journal of Insect Behavior, 18(4): 505-512.
.pdf will load immediately at http://www.cyf-kr.edu.pl/~rotofils/tofilski_ratnieks_2005.pdf

Michael H. J. Möglich and Gary D. Alpert (1979). Stone Dropping by Conomyrma bicolor (Hymenoptera: Formicidae): A New Technique of Interference Competition. Behavioral Ecology and Sociobiology, 6 (2):  105-113.
free .pdf available at Springer
http://www.springerlink.com/content/p16717229406004l/